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By 1974 the northern WCML had been electrified, and the Class 50 fleet was being transferred to BR’s Western Region to work main line passenger services out of London Paddington.
The Class 50’s 50s did not originally carry names, but in the late 1970s BR agreed to their being named after Royal Navy vessels with notable records in the First and Second World Wars.
Withdrawal of the class began in the early 1990s.
In November 2006 shareholders in Project Defiance agreed to a merger with [[The Fifty Fund]], owners of two other Class 50s and operators of a third, to form [[The Class 50 Alliance]]. Ownership of 50049 passed to the new organisation.
After the a railtour of on 18 October 2008, Defiance underwent a bogie and wheelset overhaul.The loco was taken out of traffic in 2013 following the discovery of leaking cylinder liners. A top end power unit rebuild was carried out, along with replacement of the main and ETH generators with overhauled ones.<ref>[http://www.fiftyfund.org.uk/index.php/our-locomotives/50049-defiance Fifty Fund] (retrieved 21 January 2018)</ref> 50049 completed its overhaul in April 2016 in time to work passenger services during the April 'Open House' weekend.
===50049 Defiance on the main line in preservation===
!data-sort-type="date"| Date!!Tour name !! Tour Operator!!Route !! Notes !! Reference
|-
|data-sort-value="116/9/2000" | 16 September 2000||Celtic Hoover || || ||With 50031 Hood (as D431) ||
|-
|data-sort-value="27/11/2004" | 27 November 2004||The Yorkshire Royale||Pathfinder||Cardiff-Derby (detached/reattached)-York (steam) ||With 50031 Hood and 6233 Duchess of Sutherland|| <ref name="EEOrg"/>
|data-sort-value="7/10/2017" | 7 October 2017||The Caledonian ||Pathfinder||Crewe-Glasgow ||With [[BR Class 50 50007 Hercules | 50007 Hercules]] || <ref>[https://www.youtube.com/watch?v=bKeiPbEitQU YouTube] (Retrieved 21 January 2018)</ref>
|-
|data-sort-value="14/4/2018" | 14 April 2018||Cumbrian Hoovers ||Pathfinder ||Birmingham - Settle & Carlisle -Carlisle returning via Cumbrian Coast ||Charity railtour, with [[BR Class 50 50007 Hercules | 50007 Hercules]] || <ref>[http://www.fiftyfund.org.uk/ Fifty Fund website] (Retrieved 21 January 2018)</ref>
|-
|data-sort-value="23/6/2018" | 23 June 2018|| Mazey Day Cornishman ||Pathfinder ||Tame Bridge Parkway - Penzance (50s From/to Worcester) ||With [[BR Class 50 50007 Hercules | 50007 Hercules]], the pair running as numbers 50006/11 ||
|-
|data-sort-value="5/1/2019" | 5 January 2019|| The Waverley Re-Union ||Pathfinder ||Birmingham (via Carlisle) to Edinburgh and Tweedbank ||With [[BR Class 50 50007 Hercules | 50007 Hercules]] ||
|-
|data-sort-value="23/3/2019" | 23 March 2019|| The Terminator – Phoenixed ||Pathfinder ||London Paddington to Penzance and return to Waterloo ||With [[BR Class 50 50007 Hercules | 50007 Hercules]] ||
|-
|data-sort-value="26/8/2019" | 26 August 2019|| The Cotswold Eden ||Pathfinder ||Didcot to Carlisle ||With [[BR Class 50 50007 Hercules | 50007 Hercules]] ||
|-
|}
[[Diesel Locomotives]]<br>
[[List of preservation groups]]<br>
[[SVR-based diesel locomotives visiting other events]]<br>
[[Locomotive numbering|Locomotives running under different identities]]
